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Pylons were massive, energy-based weapons designed by Red Sun. Originally built to funnel 

Titan

A pylon, close to Project:Titan.

unusable energy away from Titan, they soon doubled as defensive weapons, able to badly damage nearby Allied units. Shutting a pylon down could be acomplished in two different ways: either capture the power plant fueling it, or smash the pylon itself.

Twelve pylons surrounded the heart of Project:Titan, and were destroyed in order to overload the weapon and destroy it.

Normally, cities cannot fuel a pylon, as they amount of power they require is enormous.